Southwestern California

I had so much fun in the Peninsular Ranges last week that I headed that way again. This time, I stay much more coastal, botanizing in a three spots north of San Diego–Mission Trails Regional Park, Torrey Pines State Reserve, and Santa Rosa Plateau Ecological Reserve (confusingly, the Santa Rosa Plateau is not in the Santa Rosa mountains where I botanized last week, nor is it anywhere near the bay area city of Santa Rosa). I chose these locations to be about as different in their vegetation as possible. Mission Trails has fairly typical Coastal Sage Scrub–a chaparral habitat characterized by aromatic shrubs such as Salvia apiana (White Sage, Lamiaceae), and Mimulus aurantiacus var puniceus (Red Sticky Monkeyflower). The later is a hummingbird pollinated coastal form that grades into the more typical orange, bee-pollinated form as you move inland.

These two plants, along with the delightfully abundant Calochortus splendens (Splended Mariposa Lily), were the only species I photographed that were present at all three locations.

Calochortus_splendens

A couple bushy pink flowers in the mallow family were blooming in good numbers in the park–  Malacothamnus fasciculatus (Chapparal Mallow), and Sidalcea sparsiflora (Southern Checkerbloom). These species are in the same plant family as hibiscus and cacao (the plant from which chocolate comes).

Zeltnera venusta (Charming Centuary, Gentianaceae), with its crazy corkscrew-shaped anthers and ribbed sepals, was a nice surprise on an open hillside.

Zeltnera_venusta_2

I was the most excited to see a couple southern Clarkia species–the only all white flower in the genus, Clarkia epiloboides (Willow-Herb Farewell-to-Spring), and the rare Clarkia delecata (Delicate Farewell-to-Spring).

As their common name suggests, these are some of the last spring flowers to bloom in an area, which makes seeing them a little bittersweet. Luckily, however, I can just move my botanizing north and up in elevation.

My next stop was the Torrey Pines Reserve, just up the coast from San Diego and home of the one of the world’s rarest Pines, Pinus torreyana, a species found only here and on Santa Rosa island (Another place in California named Santa Rosa that is not near the other 3! What’s going on here?).  I managed to forget to photograph the pine trees, as I came for the coastal bluff plant community:

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If you are ever in the San Diego area, I highly recommend making a stop here–it’s San Diego’s version of Point Lobos, for bay area folks familiar with that State Park. I’ve written about coastal bluffs previously on this blog. The south coast’s version of this habitat receives much less rain and fog than the central or north coast, and subsequently has cacti, such as the Opuntia sp in the above photo and the rare Ferocactus viridescens (San Diego Barrel Cactus).

Ferocactus_viridescens_2

A few more stand outs from this location:

The showy and rare Leptosyne maritima (Sea Dahlia, Asteraceae)

Leptosyne_maritima_2

The tiny Linanthus dianthiflorus (Fringed Linanthus, Polemoniaceae)

Linanthus_dianthiflorus

And the honey-scented Piperia cooperi (Cooper’s Rein Orchid, Orchidaceae)

Piperia_cooperi

For my last stop i headed North and inland to Santa Rosa Plateau, an area with unique soils and vernal pools. Oak woodlands and grasslands dominate the landscape. This is still Southwestern California, so even here, cacti are everywhere, even growing out of the grass near a large vernal pool.

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This is the only place in the world where Brodiaea santarosae (Santa Rosa Brodiaea, Themidaceae) is found. This species is told from more common relatives by its small, triangular stamenodes (sterile stamens).

Brodiaea_santarosae

Navarretia prostrata (Pincushionplant, Polemoniaceae) was just beginning to bloom

Navarretia_prostrata

And finally, I found a couple species of one of my favorite genera, Downingia cuspida (Toothed Calicoflower, Campanulaceae) Downingia bella (Beautiful Calicoflower).

The San Joaquin Valley

The San Joaquin Valley forms the southern half of California’s vast Central Valley (Sacramento Valley sits to the north, with the San Francisco Bay delta being the dividing line). This area used to be an epic expanse of wildflower meadows and wetlands, but unfortunately, almost all of the wetlands have been drained and the meadows invaded by European annual grasses. I spent the weekend traveling to both sides of the valley–the foothills of the Greenhorn Mountains to the East and the Carrizo Plain to the West–in search of remnants of San Joaquin’s past glory. With the exception of a quick trip into the oak woodlands, I didn’t see a native tree all weekend. But I did find a few flowers. I botanized in Atriplex (Saltbush) Scrub, where plants must deal with salty, basic (high pH soils):

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and I climbed into beautiful Temblor Range (that’s Carrizo Plain in the background) :

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But mostly I hung out in the “grasslands”of the valley floor.

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Yep, a few flowers.

The color of the weekend was yellow–hills were covered in various golden members of the Aster family. Asters have flowers in dense clusters with outer “ray” and inner “disk” flowers. When you think of a sunflower or a daisy flower, you are actually thinking of a whole cluster of flowers. Many of the species look somewhat similar, and the family has its own unique set of terminology to tell them apart. You need too look at things like the phyllaries (green scales on the back of the flower cluster) and the pappus (tuft of hairs on top of the seed). Here are some of the less common asters from the weekend:

Deinandra pallida (Kern Tarweed). This species is common in the hills around Bakersfield, but is pretty much only found in Kern County. It’s just beginning to bloom.

Deinandra_pallida

Lasthenia ferrisae (Alkali Goldfields) is a specialist on salty depressions in the Central Valley. You can see the characteristic fused phyllaries in the second picture.

Monolopia congdonii (San Joaquin Woolythreads). This species does not have any ray flowers. Because the primary function of these flowers is to attract pollinators, the loss of them (which has evolved a number of times) generally indicates a self-pollinator.

Monolopia_congdonii_1

Leptosyne calliopsidea (Leafy-stemmed Coryopsis, large yellow) and Layia sp (Tidy tips, white and yellow smaller species). I messed this one up. I assumed the Layia was the common Layia platyglossa, but it turns out there are records of the very rare Layia munzii at the exact spot where I took this. You need to look at the shape of the disk flower pappus to tell them apart, and I don’t even have a close-up photo, so this will remain a mystery. I guess it still is a pretty picture though, even with unidentified plants.

Leptosybe_calliopsidea_and_layia_munzii

The weekend wasn’t all asters, though. Some additional highlights were two species of Jewelflower that were new to me–the pleasant Caulanthus anceps (Lemmon’s Mustard)

Caulanthus_anceps

And the bizarre Desert Candle (Caulanthus inflatus)

Yes, those two species are in the same genus–they are a little easier to tell apart than the Layias or the Lasthenias of the world. The weird main stem of the desert candle is hollow, so it’s a fun plant to squeeze.

Delphinium recurvatum (Byron Larkspur). A beautiful specialist in Central Valley Atriplex scrub. In this larkspur flower, the petals are actually the white parts in the middle. The sepals, which are green in most plants, are lavender here. The pointy thing in the back is the nectar spur, which contains the reward for visiting bees.

Finally, as mentioned earlier, I did briefly make it out of the valley and into the foothills. I was looking for a monkey flower species that specializes on patches of bare soil around granite outcrops. I came back with two monkeyflower species that specialize on patches of bare soil around granite outcrops! I found Mimulus congdonii (Cogndon’s Monkeyflower and Mimulus pictus (Calico Monkeyflower) hanging out right next to each other.

I was looking for the later species, as it’s both amazingly patterned and really rare, and it didn’t disappoint. But for the record, I think Mr. Joseph Congdon, a prominent 19th century Sierra botanist, has himself a very nice monkeyflower as well.